A.D. 2016, the foundations of humanity have been incinerated by the Mage King Solomon. Chaldea, a secret mages organization with the mission to preserve humanity's future, foresaw mankind's extinction in 2015. Thus commenced the operation to repair the Singularities in history caused by Holy Grails dispersed across time and space—Operation Grand Order. Using the Rayshift time travel technology, Chaldea's last master Ritsuka Fujimaru and his demi-servant Mash Kyrielight have traveled to and resolved six Singularities. Now, they depart for their most dangerous destination yet: a civilization in the Age of Gods, B.C. 2655 Mesopotamia. Ritsuka and Mash soon discover that Demonic Beasts roam the land, attacking people and towns. Amidst chaos and terror lies humanity's last defense—Uruk, a fortress city that acts as the frontline for the battle against the beasts. The battlefront is commanded by none other than King Gilgamesh, the King of Heroes, who sought aid from Heroic Spirits and took on the role of a mage to protect his city. Along with Gilgamesh and the summoned servants, Ritsuka and Mash must protect Uruk against the magical beasts' onslaught and defeat the Three Goddess Alliance who aims to eradicate humankind; all the while, a greater threat looms over Uruk, preparing for its awakening.

The year 2010 AD. Romani Archaman has been posted to Chaldea. There, he becomes the primary doctor for a young girl. Mash Kyrielight, Chaldea's second successful summoning experiment, is interested in the word "Senpai." The interaction between the two gives Mash a reason to hope. That hope becomes a wave and starts to spread. That is your story, the story of a normal "somebody." The story at the beginning of a journey that weaves the future. (Source: AniplexUS)

The first episode of "season" one was pre-aired on the Fate Project New Year's Eve TV Special 2020 program.

"This time for sure, I will kill my king with my own hands..." Jerusalem, 1273 A.D. Here, both the wandering knight, Bedivere, and Chaldea’s own expedition find the Sixth Singularity. The residents of this desolate land have been driven from their homes as three opposing powers battle for control of the land. Bedivere and company seek the help of the Mountain People to rise up against the Lion King and the Knights of the Round Table and combat their slaughter of innocents. After a deadly struggle and the loss of a dear friend, the group approaches Ozymandias, the Sun King, whose incredible power could help them turn the tables. In the halls of this great pharaoh, they desperately seek an alliance. Ozymandias, however, reveals the Lion King’s plans and refuses to assist in foiling them, instead wishing to protect his own people. Shaken to their core by the revelation of the Lion King’s true intentions, Bedivere and the Chaldean expedition decide to mount an attack on Camelot itself, hoping to put an end to the Lion King’s schemes. At last, the day of the final battle in the Holy City arrives. What are the true intentions of the Lion King, who has become so cold-blooded and diabolical? How can humanity be saved from a world where ruin has been already promised? What will become of Bedivere after shouldering the burden of a sin that can never be pardoned? (Source: Official Website)