Throughout history, humans have been at odds with Beastmen—a species capable of changing shape due to their genetic "Beast Factor." Because of this conflict, Beastmen have been forced into hiding. Anima City serves as a safe haven for these oppressed individuals to live free from human interference. During a festival celebrating the town's 10th anniversary, Michiru Kagemori, a human who suddenly turned into a tanuki, finds that Anima City is a far cry from paradise. After witnessing an explosion in the square, she is confronted by Shirou Ogami, a seemingly indestructible wolf and sworn protector of all Beastmen. As they pursue the criminals behind the bombing, the two discover that Michiru is anything but an ordinary Beastman, and look to investigate her mysterious past and uncanny abilities. Could she turn out to be the missing link between Humans and Beastmen?

Senkuu has made it his goal to bring back two million years of human achievement and revive the entirety of those turned to statues. However, one man stands in his way: Tsukasa Shishiou, who believes that only the fittest of those petrified should be revived. As the snow melts and spring approaches, Senkuu and his allies in Ishigami Village finish the preparations for their attack on the Tsukasa Empire. With a reinvented cell phone model now at their disposal, the Kingdom of Science is ready to launch its newest scheme to recruit the sizable numbers of Tsukasa's army to their side. However, it is a race against time; for every day the Kingdom of Science spends perfecting their inventions, the empire rapidly grows in number. Reuniting with old friends and gaining new allies, Senkuu and the Kingdom of Science must stop Tsukasa's forces in order to fulfill their goal of restoring humanity and all its creations. With the two sides each in pursuit of their ideal world, the Stone Wars have now begun!

After the destruction of their hometown, childhood friends Kafka Hibino and Mina Ashiro make a pact to become officers in the Defense Force—a militarized organization tasked with protecting Japan from colossal monsters known as "kaijuu." Decades later, the 32-year-old Kafka has all but given up on his dreams of heroism. Instead, he cleans up the remains of the slaughtered kaijuu after they are defeated by valiant soldiers—including Mina, who has successfully achieved their shared goal. Upon meeting his new coworker, Reno Ichikawa, Kafka faces a mirror of his past self: an ambitious young man whose one desire is to fight as a member of the Defense Force. Unfortunately, the two are soon involved in a freak encounter with a rogue kaijuu. Though Kafka demonstrates his innate heroic nature and rescues Reno from certain doom, he is left gravely injured. While both men recover in a hospital, Kafka is seemingly attacked by another one of the beasts. As a result, he gains the ability to transform into a humanoid kaijuu with the strength and powers of the massive monsters menacing Japan. Dubbed "Kaijuu No. 8" by the military, Kafka resolves to use his newfound gifts for the greater good. Tied together by mutual respect, Kafka and Reno set out to join warriors like Mina at the forefront of the Defense Force.

U.A. High School's students of Class 1-A have made it to summer break. Izuku Midoriya accompanies his mentor All Might to a celebratory superhero festival on I-Island, an isolated patch of land dedicated to researching Quirks and everything else associated with the hero business. Midoriya is granted the opportunity to meet All Might's friend Dave and Dave's daughter Melissa, two talented hero equipment engineers. He also encounters his classmates, most of whom have been given the opportunity to spend part of their summer break at the festival. However, a mysterious squad of villains infiltrates I-Island, and it is up to Midoriya and his friends to confront them, using their developing Quirks to fight off the new enemy and uncover a treacherous plot.