After dreaming about an anime she used to watch as a child, Minoa Asagaya could not forget a particularly memorable scene. However, despite her best efforts, she cannot recall the name of the show. Due to this, Minoa asks for help from her fellow classmates at Sakaneko High School. Her conversation is overheard by Arisu Kamiigusa, the most popular and wealthy girl in class who is also a hardcore otaku. Yet even with her vast knowledge, Arisu does not recognize the show. After discovering that there isn't an anime club at their school, Minoa and Arisu create the Anime Research Club, as they may obtain the answer to Minoa's mystery if they gather people who share the same interest. Thus, Minoa is exposed to a bizarre new world—the world of anime!

In the distant past, Miguel, the God of Destruction, was sealed inside the knight who ruled over light and darkness, Sturmhurt. Alongside the knight was Gestöber, who accompanied him through countless battles. In the present day, destiny causes them to reincarnate as Kabuto Hanadori and Seri Koyuki, two classmates. Their reunion should be a joyous moment, if not for the fact that these fantasied heroes are just products of Kabuto's delusions. As the fictional "Gestöber," Seri finds himself in various embarrassing situations due to Kabuto's antics that sometimes grow out of his control. Moreover, his classmate Utsugi Tsukimiya joins the fray with his absurdly accurate mind-reading abilities, slowly destroying Seri's social life. Seri tries hard to stay away from them, refusing to acknowledge their shenanigans, however with Kabuto's chuunibyou and Utsugi's unpredictability, he is only bound to be swept by the craziness coming his way.