The war between the Levamme Empire and the Amatsukami Imperium has been raging for years. In the midst of this struggle, the prince of the Levamme Empire declares his love for Juana del Moral and vows to end the war in one year, as part of his marriage proposal. When the Amatsukami catch wind of this, they assault the del Moral residence, targeting Juana's life. As a last ditch effort to bring the prince his bride, the San Maltilia Airforce hires a mercenary of mixed blood—a bestado—to fly Juana to the Levamme capital in secret. The pilot, Charles, accepts the mission...but traversing an ocean alone, into enemy territory, proves a much more dangerous ordeal than anyone could have anticipated.

Kanji Sasahara is an introverted college freshman just looking for a place to fit in. One day, he happens to stumble upon the club known as the Society for the Study of Modern Visual Culture—otherwise known as Genshiken—that serves to bring the full spectrum of otaku culture together. His first visit to the club, however, does not end well as Sasahara's pride is crushed by his senior, Harunobu Madarame, and he leaves the meeting in full denial of his otaku nature. However, after befriending club member Makoto Kousaka, who turns out to be a hardcore otaku despite his looks, Sasahara becomes more involved with club activities which include obsessing over their favorite anime, reading doujinshi, and attending conventions. There, he meets other interesting people like Kousaka's vehemently non-otaku girlfriend Saki Kasukabe, who strives to turn her boyfriend into a "normal guy." While Saki struggles to understand otaku culture and her boyfriend's love for it, Sasahara finds himself enjoying his time at Genshiken, gradually shedding any denial he once had about being an otaku and immersing himself in an otaku lifestyle.

The Society for the Study of Modern Visual Culture, otherwise known as Genshiken, is now under the charge of a more confident Sasahara. Things have changed in between semesters, and the otaku club now has a new otaku-hating member named Ogiue. Sasahara's initial goal of starting a doujin circle and selling those fan-made magazines at the next Comic Festival becomes a reality, but reality is a cruel master... who apparently crossplays. Afterward, the club is abuzz with talk about Tanaka and Ohno's relationship, which takes a hesitant step forward. (Source: Media Blasters)

Kyoto has been populated by groups of tanuki and tengu for years, living alongside humans who are oblivious to the existence of these creatures. Yasaburou Shimogamo is the third son of an influential tanuki family who spends his carefree days taking care of an old tengu, observing humans through his ability to shapeshift, and dealing with the mysterious woman named Benten. Behind the peace and tranquility, however, is a painful memory from long ago as Yasaburou's father, head of the tanuki community, was killed and eaten by a group of humans known as the Friday Fellows. Uchouten Kazoku follows the trials and tribulations of the Shimogamo brothers as they struggle to avoid their own grisly demise while coming ever closer to unraveling the truth behind their father's death.