After mustering up the courage to propose to a girl in an online game, naive otaku Hideki "Rusian" Nishimura is devastated when she flat-out rejects him. To make matters worse, the girl reveals that she is actually an older man in real life. With his dreams crushed and his heart broken, Rusian comes to an abrupt decision in the midst of his raging fit: he will never trust another girl in an online game again. Years later, Rusian is now in a guild with three other players, one of whom possesses a female avatar by the name of Ako. Ako is deeply in love with Rusian and wants to marry him. Although he entertains the possibility that she might be a guy, Rusian accepts her proposal, claiming that her gender does not matter as long as she is cute in-game. However, after a discussion between the guild members that led to all of them having an offline meeting, Rusian finds out that Ako, along with the other members, is not just a girl but also his schoolmate.

Japan's latest competitive sport, keijo, is dictated by a simple set of rules: female-only participants must stand on circular platforms floating in a pool—referred to as "lands"—with the goal being to knocking off opponents using only their breasts and butts. Despite this outlandish premise, the sport attracts millions of viewers across the country and boasts a lavish prize pool. Many aspiring athletes take up the challenge in hopes of becoming the next national champion. After graduating from high school, the lively 17-year-old Nozomi Kaminashi enters the world of keijo, hoping to bring home a fortune to her poor family. As a gifted gymnast, Nozomi quickly proves herself a tough competitor after stealing the spotlight in her debut tournament. Meeting new friends and rivals as she climbs the ranks, Nozomi discovers that the path to stardom as a keijo player is filled with intense competition that will challenge not only her body, but also her soul.