Since childhood, Hiyori Yamakawa has witnessed her father's various eccentricities, but one activity she has always loved doing with him is fly fishing—a complicated yet rewarding method of catching fish. Even after his sudden death, Hiyori still visits the seaside in her spare time, savoring the joy that her father's memento brings her. Three years later, Hiyori's mother decides to remarry and arranges a dinner with her fiancé, his daughter, and Hiyori to celebrate. Anxious and wanting to pass the time, Hiyori goes to her usual fishing spot when a rather energetic girl named Koharu Minagi suddenly appears before her. The two soon become acquainted, with Koharu taking an interest in fly fishing herself. However, their meeting is more than just a coincidence: Koharu turns out to be Hiyori's future stepsibling, which is only the beginning of treasured memories the both of them will reel in as they foster their sisterhood.

Ten years have passed since Nagomu Irino left his home to pursue his dream that ultimately failed. He finally decides to return after hearing that his father—owner of the Kyoto sweets store Ryokushou—has been hospitalized. Worrying that no one would inherit the shop if his father passes away, Nagomu prepares himself to embrace his family's legacy and the art of making sweets⁠. Unsurprisingly, Nagomu discovers that his father has already chosen a different successor—a 10-year-old girl named Itsuka Yukihira, who was abandoned at Ryokushou by her father for reasons unknown. While she has since become a part of the family and is now the shop's poster girl, Itsuka still longs to see her father and follows all possible clues that may lead her to him. Sympathizing with Itsuka's situation, Nagomu's mother asks Nagomu to act as Itsuka's father, hoping that Itsuka will open up to him and relieve her pain, even if just a little. Starting on bad terms, Itsuka and Nagomu gradually learn more about each other, realizing that they are more similar than they had thought. Connected by their mutual love for both Ryokushou and its confections, their relationship as child and father figure begins to make lives around them a little bit sweeter.

In the kingdom of Highland, fairies lived freely until humans betrayed them. Once a fairy has been stripped of a wing, whoever possesses it holds the fairy's life in their hands. Taking advantage of this, humans enslaved fairies, but 15-year-old Ann Halford longs to see the day when fairies and humans are treated as equals. The aspiring confectioner dreams of becoming a Silver Sugar Master just like her late mother. In order to achieve this lofty goal, Ann sets out for the capital of Lewiston, home to the annual Royal Candy Fair. However, to be bestowed the elusive title, Ann's confection crafted from silver sugar must be recognized with a royal medal at the event. Ann hires a sharp-tongued warrior fairy, Shall Fen Shall, to be her bodyguard for the perilous journey ahead. After saving him from his cruel fairy trader, she promises to return Shall's wing to him once they reach the capital, much to his confusion. But as the pair travel further along, the time they spend together grows, and they learn more about each other and their true desires.

Apprentice mage Chise Hatori is invited to enroll at the College, a prestigious learning institution for sorcerers, to examine and look for a way to remove the curses she bears. Despite the reluctance of her groom, Elias Ainsworth, Chise accepts the proposal, as she believes attending the school might help her minimize her self-sacrificing tendencies. From the get-go, Chise grabs the attention of her classmates and professors alike, who have never seen a mage in action before. However, there is a sinister plot brewing behind the College's back, and the young mage will have to determine who is friend or foe in order to put a stop to it.